Its source is located in the Lusatian Hills (Lausitzer Bergland) on the Czech border. After leaving the hills, the river crosses the old city of Bautzen/Budyin, the centre of the Sorbs in Upper Lusatia. Further north the river enters the Spreewald, a large wetlands area, which belongs to Lower Lusatia. In its final portion the river runs through the city centre of Berlin to join the River Havel in Spandau, a western quarter of Berlin.

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